Ray Romano
Manny (voice)
John Leguizamo
Sid (voice)
Denis Leary
Diego (voice)
Adam Devine
Julian (voice)
Jesse Tyler Ferguson
Shangri Llama (voice)
Max Greenfield
Roger (voice)
Jessie J
Brooke (voice)
Nick Offerman
Gavin (voice)
Keke Palmer
Peaches (voice)
Josh Peck
Eddie (voice)
Simon Pegg
Buck / Pythagorus Buck / Robo Buck (voice)
Seann William Scott
Crash (voice)
Wanda Sykes
Granny (voice)
Jennifer Lopez
Shira (voice)
Queen Latifah
Ellie (voice)
Stephanie Beatriz
Gertie (voice)
Neil deGrasse Tyson
Neil deBuck Weasel (voice)
Lilly Singh
Bubbles and Misty (voice)
Michael Strahan
Teddy (voice)
Melissa Rauch
Francine (voice)
Chris Wedge
Scrat (voice)
Robert Cardone
Freaky Bobby (voice)
Byron Thames
Additional Voices (voice)
Rif Hutton
Additional Voices (voice)
Chris Campbell
Additional Voices (voice)
Holly Dorff
Additional Voices (voice)
Moosie Drier
Additional Voices (voice)
Jason Fricchione
Additional Voices (voice)
Jackie Gonneau
Additional Voices (voice)
Hope Levy
Additional Voices (voice)
Terence Mathews
Additional Voices (voice)
Edie Mirman
Additional Voices (voice)
Juan Pope
Additional Voices (voice)
Ruth Zalduondo
Additional Voices (voice)
David Zyler
Additional Voices (voice)
Ashley Peldon
Additional Voices (voice)
Carlos Ponce
Mariachi Beaver (voice)
Michael Thurmeier
Gravedigger Beaver / Party Molehog (voice)
Randy Thom
Hyrax Minister (voice)
Jorge Lucas
Grandpa Molehog (voice)
James Palumbo
Panicked Start (voice)